What is the opposite of :hover (on mouse leave)?

Question

Is there any way to do the opposite of :hover using only CSS? As in: if :hover is on Mouse Enter, is there a CSS equivalent to on Mouse Leave?

Example:

I have a HTML menu using list items. When I hover one of the items, there is a CSS color animation from #999 to black. How can I create the opposite effect when the mouse leaves the item area, with an animation from black to #999?

jsFiddle

(Have in mind that I do not wish to answer only this example, but the entire "opposite of :hover" issue.)

Answer 1

If I understand correctly you could do the same thing by moving your transitions to the link rather than the hover state:

ul li a {
    color:#999;       
    transition: color 0.5s linear; /* vendorless fallback */
    -o-transition: color 0.5s linear; /* opera */
    -ms-transition: color 0.5s linear; /* IE 10 */
    -moz-transition: color 0.5s linear; /* Firefox */
    -webkit-transition: color 0.5s linear; /*safari and chrome */
}

ul li a:hover {
    color:black;
    cursor: pointer;
}

http://jsfiddle.net/spacebeers/sELKu/3/

The definition of hover is:

The :hover selector is used to select elements when you mouse over them.

By that definition the opposite of hover is any point at which the mouse is not over it. Someone far smarter than me has done this article, setting different transitions on both states - http://css-tricks.com/different-transitions-for-hover-on-hover-off/

#thing {
   padding: 10px;
   border-radius: 5px;

  /* HOVER OFF */
   -webkit-transition: padding 2s;
}

#thing:hover {
   padding: 20px;
   border-radius: 15px;

  /* HOVER ON */
   -webkit-transition: border-radius 2s;
}